Placolith (or modified placolith) coccoliths with shields formed from elements that display little or no imbrication and equal development of V and R crystal units, i.e. one shield (distal) formed from V units, the other from R units (proximal). LM images are typically low birefringence but high relief in phase contrast. Shield elements are typically joined along radial sutures, but these may curve or kink. Element curvature is consistently dextrogyre (veeing anticlockwise) and obliquity is broadly dextral (clockwise) in the distal shield, and the same in the proximal shield when viewed proximally.