Family CERATOLITHACEAE Norris 1965
Family characterised by ornate horseshoe-shaped nannoliths formed from a single calcite crystal unit, termed ceratoliths. Includes the extant genus Ceratolithus and extinct genus Amaurolithus, which have similar shape but different crystallographic orientations.
Taxa included:
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| A. amplificus |
A. delicatus |
A. primus |
A. tricorniculatus | C. armatus |
C. atlanticus |
C. cristatus |
| Amaurolithus - dark in xpl |
Ceratolithus - bright in xpl |
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Life cycle: Classic observations of Norris (1971) showed that typically a single ceratolith was wrapped around the cell and that beyond the ceratolith a large coccosphere of hoop-shaped coccoliths sometimes occurs. These large coccospheres can contain up to four cells each with ceratoliths.
More recently, Alcober & Jordan (1997) observed C. cristatus hoop-shaped coccoliths inside coccospheres of "Neosphaera coccolithomorpha" planoliths, suggesting that ceratoliths, planoliths and hoop coccoliths may form during alternate phases of a complex life-cycle. These observations have been strongly confirmed by Young et al. (1998), Cros et al. (2000) and Sprengel & Young (2000). The “Neosphaera” planoliths show typical heterococcolith features, hence a likely hypothesis is that the ceratolith stage is equivalent to the holococcolith stage in other taxa, and so haploid (see also discussion in Young et al. 2005). Molecular genetic data and culture observations not yet available.
Neogene: Ceratolithaceae
Nannoliths - informal grouping. Used here in the sense of Young & Bown (1987)
Description - the following groups of Neogene nannofossils are thought to be related to coccolithophores, but produce structures which are neither heterococcoliths nor holococcoliths.
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| Braarudosphaeraceae | Ceratolithaceae | Discoasteraceae | Sphenolithaceae | Triquetrorhabdulaceae |
The term nannolith has been used, especially by palaeontologists, as a convenient term for structures about the same size as coccoliths and occurring with coccoliths, but lacking definite coccolith affinities. In the modern nannoflora, there are fewer groups of cryptic origins, and the term has been less widely used. However, it is useful for calcareous structures that are thought to be formed by haptophytes, but probably by a different biomineralisation process to either heterococcoliths or holococcoliths (Young et al. 1999).
NB The first use of the term in this sense appears to have been by Haq (1978, in Introduction to Marine Micropaleontology), and this was followed by Perch-Nielsen (1985, in Plankton Stratigraphy). By contrast, Aubry (1988 et seq., Handbook of Calcareous Nannofossils) places most of these groups in the Ortholithae.