Amaurolithus tricorniculatus (Gartner, 1967) Gartner and Bukry, 1975 [Ceratolithus]
Description: Right arm extended into apical spur.
Remarks:
Original description:
Variant: A. bizzarus (Bukry, 1973) Gartner and Bukry, 1975 [Ceratolithus] - an ornate variant of A. tricorniculatus, with additional spines. (NB In the original publication the species-name was spelt bizzarus, so it cannot be corrected to bizarrus, even though that would appear a more logical spelling).
Neogene: Amaurolithus; Amaurolithus tricorniculatus; Ceratolithus
Amaurolithus primus (Bukry and Percival, 1971) Gartner and Bukry, 1975 [Ceratolithus]
Description: Simple form with short arms, broad arch and weakly developed keels.
Remarks:
Original description:
Neogene: Amaurolithus; Amaurolithus primus; Ceratolithus
Amaurolithus delicatus Gartner and Bukry, 1975
Description: Simple horseshoe-shaped amauroliths. Grades into the other morphotypes, especially A. primus and A. tricorniculatus.
Remarks:
Original description:
Variant: A. ninae Perch-Nielsen, 1977 - like A. delicatus but with well developed apical region.
Neogene: Amaurolithus; Amaurolithus delicatus
Amaurolithus amplificus (Bukry and Percival, 1971) Gartner and Bukry, 1975 [Ceratolithus]
Description: Like A. primus but angular; right arm with rodded keel and longer than left.
Original description:
Alternative combination:
Nicklithus amplificus (Bukry and Percival, 1971) Raffi Backman & Rio 1998.
Raffi et l. (1998) presented evidence that A. primus and A. amplificus evloved independantly from Triquetrotrhabdulus and so proposed a new genus, Nicklithus (in honour of N Shackleton) for A. amplificus. I have followed the conventional nomenclature here, but comments on this are welcome (JRY).
Synonym:
C. dentatus Bukry, 1973.
Neogene: Amaurolithus; Amaurolithus amplificus; Ceratolithus
Genus Amaurolithus Gartner and Bukry, 1975
Description: Ceratoliths non-birefringent or only weakly birefringent. The c-axis is perpendicular to the plane of horseshoe, but spines on the surface may result it in being tilted from the horizontal and so showing some birefringence, but birefringence is lower than in Ceratolithus and extinction is rarely exactly in line with axis of the ceratolth.
In the SEM Amaurolithus ceratoliths (sometimes termed amauroliths) usually come to rest with the more ornamented surface upward, in this orientation the left hand arm is distinctly curved whilst the right hand arm is relatively straight, but more heavily ornamented.
Remarks: Raffi et al. (1998) documented likely evolutionary relationships between Ceratolithus and the fossil genera Amaurolithus and Triquetrorhabdulus. They presented evidence that A. delicatus, A. amplificus and C. acutus all independently evolved from Triquetrorhabdulus. Since this indicated that Amaurolithus was polypyhletic they proposed dividing the genus in two; Amaurolithus and Nicklithus.
Taxa included:
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| A. primus | A. delicatus | A. tricorniculatus | A. amplificus |
| Amaurolithus | Nicklithus | ||
Original description:
Neogene: Amaurolithus; Ceratolithus