Emiliania huxleyi

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Cenozoic Taxa

Gephyrocapsa muellerae

Gephyrocapsa muellerae Bréhéret 1978

Description: Intermediate size species of Gephyrocapsa. Coccoliths with bridge at low angle to long axis, central area rather small.

Remarks: A cold water species, dominates Gephyrocapsa assemblages at high laltitudes. In older literature on living nannoplankton G. muellerae is often refered to as G. caribbeanica.

Size: Coccospheres, 5 to 9 µm; coccoliths, 3 to 4 µm long.

G, muellerae (modern)
G. muellerae (modern)

Geological Time Data
Geological Time Periods: 
Biblio Reference: 
Young, JR.  1998.  Neogene. Calcareous nannofossil biostratigraphy. :225-265.
Notes on occurrence information: 
Common from MIS7 to MIS1 (Holocene). Frequently dominates higher latitude assemblages in MIS4-6, then is replaced by Emiliania huxleyi.

Emiliania huxleyi

Emiliania huxleyi (Lohmann 1902) Hay & Mohler, in Hay et al. 1967

Description: Ubiquitous species, often forming blooms. Coccospheres often with multiple layers of coccoliths.

Remarks: The open structure of E. huxleyi coccoliths makes them highly vulnerable to both overgrowth and etching.

Original description:

Cat050156 copy.jpgCat050157 copy.jpg

Ehux173-44NN21.JPGEhux173-04NN21.JPGE_hux_218-01a.JPGE_hux_181-61.JPG

Variability of extant E. huxleyi


Well-established morphotypes
: Young & Westbroek (1991) distinguished four types (A, B, C & corona) based on heterococcolith morphology, see also van Bleijswijk et al. (1991) and Young (1994). Medlin et al. (1996) consider these types should be regarded as separate varieties, their recommended names are given in brackets, (). Additonal recently recognised are type R and type B/C (Young et al. 2003). It should be noted that further variation is seen in degree of calcification, leading to variable central area closure and slitting of proximal shield.

E_hux_113-19.JPG Type A (huxleyi) liths medium -sized (3-4 µm), distal shield elements robust, central area elements curved
E_hux_111-22.JPG
Type B (pujosiae) liths large (3.5-5 µm), distal shield elements delicate, central area
elements irregular laths. Proximal shield often wider than distal
shield. Similar large E. huxleyi forms have been recognised in the fossil record and are regarded by some authors as cold-water forms.
E_hux_114-39.JPG Type C (kleijneae) liths small (2.5-3.5 µm), distal shield elements delicate, central area open or covered by thin plate
E_hux_131-07.JPG Type B/C Liths medium-sized, (3-4 µm), distal shield elements delicate, i.e. similar in morphology to types B & C, but intermediate in size. This form often dominates assemblages in the Southern Ocean (Hockfield 2000; Findlay & Giraudeau 2000, referred to as type C)
E_hux_167-17.JPG E. huxleyi var. corona (Okada & McIntyre 1977) Jordan & Young 1990 Like type A, but with inner tube cycle forming discontinuous elevated crown around central area
E_hux_186-11.JPG Type R Like type A, but with heavily calcified shield elements, indeed slits often closed giving a Reticulofenestra-like appearance (hence designation as type R). Several specimens of this form have been isolated from the S.W. Pacific and they have maintained their distinctive morphology for several years indicating that this is stable genotypic variation (Probert, Young unpubl. obs). Similar morphotypes were observed in the same area by Nishida (1979) and Burns
(1977)
Geological Time Data
Geological Time Periods: 
Biblio Reference: 
Young, JR.  1998.  Neogene. Calcareous nannofossil biostratigraphy. :225-265.

Emiliania

Emiliania Hay & Mohler in Hay et al. 1967

Description: Coccoliths with slits between all distal shield, and some proximal shield, elements; no bridge.

Remarks

Type: Emiliania huxleyi.

Geological Time Data
Geological Time Periods: 
Biblio Reference: 
Young, JR.  1998.  Neogene. Calcareous nannofossil biostratigraphy. :225-265.
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