Description: Conical or cylindrical nannoliths consisting of 10-30 wedge-shaped, radially-arranged elements. Typically have a concave end that is assumed to be proximal and a convex or pointed end that is considered distal. Outer surface is often covered by depressions (fenestrae). Typically seen in diagnostic side view.

Remarks: Apparently distinct distal cycles are developed in some species but these are probably formed by kinking of the elements, rather than being separate crystal-units.

Range: Predominantly upper Paleocene group (NP5 to NP10).